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As an academic discipline emerged with the modern synthesis in the 1930s and 1940s. In the 1930s the study of animal behavior (ethology) emerged with the work of the Dutch biologist Nikolaas Tinbergen and the Austrian biologists Konrad Lorenz and Karl von Frisch. Hamilton’s (1964) papers on inclusive fitness and ‘s (1972) theories on reciprocity and parental investment helped to establish evolutionary thinking in psychology and the other social sciences.

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